Author: آمنه غلامی 
Astragalus dictyolobus C.A. Meyer ex Bunge
Mohammad Dadmand, Masoumeh Ramezani Yeganeh, Mahnaz Heydari Rikan and Shariar Yeilaghi
Natural Resources Gene Bank, Research Institute of Forests and Rangelands, Agricultural Research, Education and Extension Organization, Tehran, Iran.
Uploaded at December 2025
Scientific editor: Parvin Salehi Shanjani
Seed Storage
The storage behavior of this species is orthodox (desiccation-tolerant seeds), dry to 15-20% eRH, and store at -20°C.
Germination requirements
The seeds have physical-physiological dormancy and this is related to seed coat and inhibitors. To remove the seed coat, different methods of scarification can be used (sandpaper, sulfuric acid, hot water, etc.). To break dormancy, stratification with pre-chilling (for 14 days), ultrasonic waves, and hormones (Gibberellic acid 100 ppm) are efficient. The best method for breaking dormancy is rubbing the seeds with sandpaper and moist chilling (14 days at 4 °C). Then germinate under a constant temperature of 20 ºC or alternating temperature 25 ºC (16 hours of light) and 15 ºC (8 hours of dark).
Morphology
Fruit type: Pod
Fruit shape: Perfect bilocular (divided into two separate sections), small, pubescent, valves yellowish green, reticulated, stalkless, enclosed within a calyx, ventral part convex, grooved, dorsal part flat
Seed shape: Reniform, compressed, hilum sunken, with pale halo. Surface wrinkled, glabrous, slightly lustrous, dark green, often faint spotted
Seed size: Length 4.19 to 4.95 mm, width 2.72 to 3.39 mm in an accession from khoy, Azarbayjan, with 1000 seeds weight 30.4 g, which was collected and identified in 2018 by Mahnaz Heydari Rikan and Shariar Yeilaghi and is stored in the Natural Resources Gene Bank of Iran with an accession number 47199
Average 1000 seed weight: 25 to 30.4 g
Biological features
Life form: Perennial, woody at the base, pubescent. Stems erect, 10 to 50 cm long
Inflorescence, flower type, and pollination mechanism: Inflorescence raceme, many-flowered; flowers hermaphrodite, zygomorphic, papilionaceous, corolla (composed of banner: upper large petal, wings: two petals adjacent to the banner, and keel: two bottom petals) yellow to pale brown, wings as long as the keel. Plant monoecious, cross-pollinated. Pollination is carried out by insects and wind.
Time of flowering: Spring to early summer
Time of seed ripening: Summer
Seed dispersal methods: Seed weight and animals
Geographical distribution: Endemic to Iran. Typus from Khoy, Azarbayjan
Distribution in Iran: Northwest and West. Mostly in province Azarbayjan, zanjan and Hamedan
Taxonomy
Family: Fabaceae
Genus: Astragalus
Species: A. dictyolobus.
Persian name: Gavan
References and Links
Black, M. and Bewley, J.D. 2000. Seed technology and its biological basis. Sheffield Academic Press, 1-419 pp.
Bojňanský, V. and Fargašová, A. 2007. Atlas of seeds and fruits of central and East-European flora: the Carpathian Mountains region. Springer, Dordrecht, The Netherlands, 1046 pp.
Ghahraman, A. Flore of Iran. Research Institute of Forests and Rangelands with University of Tehran Publication. https://floraofiran.net/index.php.
Javadi, H. 2021. Seed: Processing and Quality tests. Agricultural Education & Extension Publishers, 175 pp.
Maassoumi, A.A. 2003. Flora of Iran, Papilionaceae (Astragalus I). Institute of Forests and Rangelands Press, Tehran, Iran, Publication No. 43, 368 pp.
Mozaffarian, V.A. 1998. Dictionary of Iranian plants names. Farhange Moaser Publisher, Tehran, 671 pp.
Mozaffarian, V.A. 2012. A Pictorial dictionary of botany. Farhange Moaser Publisher, Tehran, 1088 pp.
Sharma, O.P. 2009. Plant taxonomy 2E. Tata McGraw-Hill Education, New Delh
