Author: آمنه غلامی 
Lepidium buschianum Al- Shahbaz
Masoumeh Izadpanah, Amene Gholami, Mohammadtaghi Feyzi and Azadeh Akhvan
Natural Resources Gene Bank, Research Institute of Forests and Rangelands, Agricultural Research, Education and Extension Organization, Tehran, Iran.
Uploaded at february 2026
Scientific editor: Parvin Salehi Shanjani
Seed Storage
The storage behavior of this species is orthodox (desiccation-tolerant seeds), dry to 15 to 20% eRH, and store at -20°C.
Germination requirements
The seeds of this species are not dormant. wet seeds mucilaginous Sterilize seeds before placing them on wet germination paper. Place seeds for two weeks at 4 ºC to germinate uniformly. Then germinate under an alternating temperature regime of 15/20 ºC (the higher temperature for 16 hours in light and the lower one for eight hours in dark).
Morphology
Fruit type: Silicule
Fruit shape: Obovate, often oblique, one seeded, valves strongly swollen, with a single vein
Fruit size: Length 5.3 to 6.33 mm, width 3.82 to 4.36 mm, pedicel 10 to 13 mm
Seed shape: Obovate, not compressed, radicular furrow distinct. Surface with indistinct ridges, glabrous, dull, brown
Seed size: Length 3.4 to 5.25 mm, width 2.75 to 3.86 mm in an accession from Fereydunshahr, Isfahan with 1000 seeds weight 17.01 g, which was collected and identified in 2019 by Mohammadtaghi Feyzi and Azadeh Akhvan, and is stored in the Natural Resources Gene Bank of Iran with the accession number 48266
Average 1000 seed weight: 17 to 21 g
Biological features
Life form: Herbaceous, perennial, glabrous. Stems erect, furrowed, branched in the upper part, up to 1 m long
Inflorescence, flower type, and pollination mechanism: Inflorescence raceme; flowers hermaphrodite, regular; polysepalous, sepals oval or elliptic – orbicular, deciduous, with prominent veins, color green with white margin; polypetalous, petals broadly-ovate or almost orbicular, with prominent veins and short linear unguis, 4 to 5 mm long, petals twice the size of the sepals, white. Plant monoecious, cross-pollinated. Pollination is carried out by insects and wind.
Time of flowering: June
Time of seed ripening: August
Seed dispersal methods: Seed weight
Geographical distribution: Endemic to Iran. Typus from Azerbaijan
Distribution in Iran: West and Center. Mostly in provinces Azerbaijan, Hamedan, Lorestan, and Tehran
Taxonomy
Family: Brassicaceae
Genus: Lepidium
Species: L. buschianum
Persian name: Tertizak-e kuhi
References and Links
Assadi, M. Sajedi, S. Fakhr Ranjberi, H. Mirzadeh Vaghefi, S. Moazzeni, H. Khodashenas, M. Khosravi, A. R. Hatami, A. Mehrnia, M. Kaffash, Sh. Heidarnia, N. Sheidai, M. Heidari Rikan, M. Kavousi, K. Sonboli, A. Veiskarami, Gh. And Aminian, F. 2017. Flora of Iran, Brassicaceae, No.143. Research Institute of Forests and Rangelands Press, Tehran, Iran, 954 pp.
Black, M. and Bewley, J.D. 2000. Seed technology and its biological basis. Sheffield Academic Press, 419 pp.
Bojňanský, V. and Fargašová, A. 2007. Atlas of seeds and fruits of central and East-European flora: The Carpathian Mountains region. Springer, Dordrecht, The Netherlands, 1046 pp.
Ghahraman, A. Flore of Iran. Research Institute of Forests and Rangelands with University of Tehran Publication. https://floraofiran.net/index.php.
Mozaffarian, V.A. 1998. Dictionary of Iranian plants names. Farhange Moaser Publisher, Tehran, 671 pp.
Mozaffarian, V.A. 2012. A Pictorial dictionary of botany. Farhange Moaser Publisher, Tehran, 1088 pp.
Sharma, O.P. 2009. Plant taxonomy 2E. Tata McGraw-Hill Education, New Delhi, 563 pp.
